The article is devoted to the problem of non-linguistic students’ English fluency and proficiency development through information and communication technologies (ICT), in particular with Internet-based project activity. As a target language in the University training system of future specialists, the English teaching course aims to provide students with an opportunity to acquire communicative and professional foreign language competence. Thus, the aim of the study is to identify the effective ways of the English fluency skills development through ICT in the university course. The article analyzes the key concept of “information and communication technology” from the point of view of Russian and foreign researches. The authors view this term as a set of tools and methods for converting information data to obtain a new quality information product. Due to the global society informatization, the revision of English teaching objectives and content as well as the search for new ways to improve teaching methods the relevance of ICT application is proved. A list of basic criteria for teachers to evaluate Internet sites is developed. Based on the main characteristics of the educational Internet project the authors determine the conditions and advantages of mastering students’ English fluency and proficiency. The paper gives specific practical recommendations on how to organize Internet-based project activities. The authors conclude that ICT is a powerful and effective resource to improve the quality of English teaching in the University course due to authentic material introduction into the educational process and creating a real environment for foreign language communication.